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Phloem

anatomy

FLOW-em

Simple Definition

The inner bark tissue that carries sugars and nutrients from the leaves down to the roots and other parts of the tree.

Technical Definition

A complex vascular tissue composed of sieve elements, companion cells, phloem parenchyma, and phloem fibers that transports photosynthates (primarily sucrose) from source organs to sink organs via translocation.

📚 Etymology

From Greek 'phloios' (bark), referring to its location in the bark region of the stem.

What is Phloem?

Phloem is the tree's food-delivery system. While xylem carries water upward from the roots, phloem carries sugars and nutrients downward from the leaves to every living part of the tree — roots, growing tips, flowers, and fruits.

How It Works

Key Components

  1. Sieve tubes: Long, connected cells that form the pipeline for sugar transport.
  2. Companion cells: Metabolic helpers that keep sieve tubes alive and functioning.
  3. Phloem parenchyma: Storage cells for starch and other nutrients.
  4. Phloem fibers: Structural support (bast fibers used in traditional cordage).

Transport Mechanism

Sugars are loaded into sieve tubes at the leaves, creating osmotic pressure that drives flow toward areas of lower sugar concentration (roots, growing points).

Location in the Trunk

Phloem sits just outside the cambium, forming the inner bark. This places it in a vulnerable position — any damage that rings the trunk (girdling) cuts the phloem and starves the roots.

Costa Rican Examples

Guapinol (Hymenaea courbaril)

Thick, resinous phloem produces copious amber-like resin when wounded.

Burío (Heliocarpus appendiculatus)

Strong phloem fibers (bast) are traditionally extracted for making rope and cordage.

Practical Significance

  • Girdling kills trees: Removing bark in a ring around the trunk cuts the phloem and starves the roots.
  • Latex production: Many tropical trees (rubber, chicle) produce latex in specialized phloem cells.
  • Traditional fiber: Bast fibers from phloem have been used for millennia in rope-making.

🌳 Example Species

Cedro Amargo

Cedrela odorata

The Cedro Amargo is one of the most valuable timber trees in the Americas, known for its fragrant, rot-resistant wood used in fine furniture, cigar boxes, and traditional crafts. Despite heavy exploitation, it remains an important species in Costa Rican forests.

Espavel

Anacardium excelsum

The Espavel is a majestic emergent tree of Costa Rica's riparian forests, reaching heights of 50 meters. A close relative of the cashew, it produces edible nuts prized by wildlife and humans alike.

Guapinol

Hymenaea courbaril

The Guapinol, or Jatobá, is a magnificent legume tree producing amber-like resin, hard durable wood, and stinky but nutritious seed pods that have sustained humans for millennia. Its resin preserves ancient insects as prehistoric amber.

🔗 Related Terms

Bark

The protective outer covering of a tree's trunk, branches, and roots.

Cambium

A thin layer of actively dividing cells between the bark and the wood that makes a tree grow thicker.

Sapwood

The outer, living wood of a tree trunk that transports water and nutrients from roots to leaves.

Xylem

The woody tissue inside a tree that carries water and minerals upward from the roots to the leaves.

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